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  <front>
    <journal-meta />
    <article-meta>
      <title-group>
        <article-title>Assessment of the Tourist Carrying Capacity in the Manglaralto River-Aquifer System, Santa Elena (Ecuador): A UNESCO Ecohydrology Demonstration Site</article-title>
      </title-group>
      <contrib-group>
        <contrib contrib-type="author">
          <string-name>María Jaya-Montalvo</string-name>
          <xref ref-type="aff" rid="aff0">0</xref>
        </contrib>
        <contrib contrib-type="author">
          <string-name>Lady Soto-Navarrete</string-name>
          <xref ref-type="aff" rid="aff0">0</xref>
          <xref ref-type="aff" rid="aff2">2</xref>
        </contrib>
        <contrib contrib-type="author">
          <string-name>Vicky Arteaga</string-name>
          <email>vicky181991@hotmail.com</email>
          <xref ref-type="aff" rid="aff3">3</xref>
        </contrib>
        <contrib contrib-type="author">
          <string-name>Josep Mata-Perelló</string-name>
          <xref ref-type="aff" rid="aff1">1</xref>
        </contrib>
        <contrib contrib-type="author">
          <string-name>Paúl Carrión-Mero</string-name>
          <xref ref-type="aff" rid="aff0">0</xref>
        </contrib>
        <aff id="aff0">
          <label>0</label>
          <institution>ESPOL Polytechnic University</institution>
          ,
          <addr-line>Guayaquil</addr-line>
          ,
          <country country="EC">Ecuador</country>
        </aff>
        <aff id="aff1">
          <label>1</label>
          <institution>Polytechnic University of Catalonia</institution>
          ,
          <addr-line>Barcelona</addr-line>
          ,
          <country country="ES">Spain</country>
        </aff>
        <aff id="aff2">
          <label>2</label>
          <institution>Universitat Rovira i Virgili</institution>
          ,
          <addr-line>Tarragona</addr-line>
          ,
          <country country="ES">Spain</country>
        </aff>
        <aff id="aff3">
          <label>3</label>
          <institution>University of Guayaquil</institution>
          ,
          <addr-line>Guayaquil</addr-line>
          ,
          <country country="EC">Ecuador</country>
        </aff>
      </contrib-group>
      <fpage>277</fpage>
      <lpage>288</lpage>
      <abstract>
        <p>The UNESCO-sponsored ecohydrology demonstration sites function as natural spaces designed for water purification and nutrient reduction, promoting sustainable management based on both technical and ancestral ecological processes. The unique characteristics of these sites have attracted the interest of visitors, researchers, and students interested in ecotourism, environmental education, and conservation. Assessing tourism carrying capacity plays a key role in ensuring the ecological, social, and cultural sustainability of such sites. This study aims to determine the tourist carrying capacity of the Manglaralto River-Aquifer System, identifying interactions between visitors and water resources using analytical tools, to promote sustainable and responsible management of the water resources. The methodology involved applying a cause-and-efect matrix, which allowed for the visualisation of six negative components, with magnitudes ranging from -3.5 to -9, that afect the interaction between environmental quality and tourism performance. This assessment allowed for the definition of management strategies, such as reforestation, signage, and community participation. The approach of these measures integrates ecohydrological practices as an educational tourism element, promoting environmental awareness and positioning Manglaralto as a replicable model for marine coastal systems sensitive to anthropogenic activity.</p>
      </abstract>
      <kwd-group>
        <kwd>eol&gt;Geosites</kwd>
        <kwd>Ecology</kwd>
        <kwd>Aquifers</kwd>
        <kwd>Sustainability</kwd>
      </kwd-group>
    </article-meta>
  </front>
  <body>
    <sec id="sec-1">
      <title>1. Introduction</title>
      <p>
        For decades, people have taken advantage of places with singular natural landscapes to engage in
tourism, causing significant modifications to the ecosystem. The development of tourism is usually
supported using land for the construction of infrastructure, the creation of regulations related to
overaccommodation and construction, and the promotion of historical, cultural, and natural attractions
[
        <xref ref-type="bibr" rid="ref1">1</xref>
        ]. Marine-coastal tourism is growing daily and is key to the economic reports of each sector [
        <xref ref-type="bibr" rid="ref2">2</xref>
        ].
Mark [
        <xref ref-type="bibr" rid="ref3">3</xref>
        ], defined marine tourism as recreational activities carried out on a trip that involve the marine
environment. On many occasions, tourists are unaware of the conservation of marine coastal fauna and
associated ecosystems, such as beaches, plains, tidal flats, dunes, and estuaries [
        <xref ref-type="bibr" rid="ref4 ref5">4, 5</xref>
        ].
      </p>
      <p>
        One way to mitigate visitor damage in tourist areas is through carrying capacity, a definition first
introduced in 1921 [
        <xref ref-type="bibr" rid="ref6 ref7">6, 7</xref>
        ], and described as the most significant capacity of an environment to resist
environmental changes. Carrying capacity was mainly used in the field of ecology to determine
the number of animals or plants in a specific area [
        <xref ref-type="bibr" rid="ref8">8</xref>
        ]. Later, it was modified and conceptualised
as the assumption of the area available for tourism and the number of visitors that the place can
accommodate without altering its ecosystem, considering the physical, biological, environmental
factors and characteristics of the place [
        <xref ref-type="bibr" rid="ref10 ref9">9, 10</xref>
        ]. There are two approaches to tourism carrying capacity:
the first covers the destination’s willingness to accommodate tourists before damage is caused to the
ecosystem, and the second is based on the negative experience of the tourist’s perception of the poor
conditions of the environment, making the place less satisfactory and attractive [
        <xref ref-type="bibr" rid="ref11">11</xref>
        ].
      </p>
      <p>
        In recent years, the need to preserve coastal tourism sites has led to tourism development plans
being adjusted to include broader environmental and sociocultural concerns [
        <xref ref-type="bibr" rid="ref12 ref13">12, 13</xref>
        ]. Since 2011, the
United Nations Educational, Scientific and Cultural Organisation (UNESCO) through the International
Hydrological Programme (IHP) has promoted the creation of demonstration sites in various parts of the
world, implementing ecohydrological solutions in watersheds at diferent scales [
        <xref ref-type="bibr" rid="ref14">14</xref>
        ]. The demonstration
sites aim to showcase ecohydrology as a transdisciplinary science. In 2001, the IHP supported initiatives
that achieved scientific progress and fostered transparency of knowledge, promoting education. To this
end, criteria have been defined to identify places where innovative and sustainable water management
is implemented [
        <xref ref-type="bibr" rid="ref15">15</xref>
        ].
      </p>
      <p>
        The concept of ecohydrology is represented by the approach to the restoration and sustainability
of water resources, serving as an additional tool to control the ecological degradation of water and
surface processes [
        <xref ref-type="bibr" rid="ref5">5</xref>
        ]. Within the framework of water sustainability, ecohydrology relates hydrology to
environmental and biotic processes [
        <xref ref-type="bibr" rid="ref16">16</xref>
        ]. This approach seeks to understand and take advantage of the
functions of these processes to achieve a correct balance in ecosystems, especially those that have been
disturbed by humans [
        <xref ref-type="bibr" rid="ref17">17</xref>
        ]. In coastal areas, ecohydrology is key to the sustainability of the ecosystem,
as the process of interaction between freshwater and saltwater mitigates the impacts of coastal erosion
and saline pollution [
        <xref ref-type="bibr" rid="ref18">18</xref>
        ].
      </p>
      <p>
        Coastal ecohydrology focuses on the restoration of the carrying capacity of estuarine and coastal
areas, allowing the concept of management, harmonisation, and solutions to problems in basins [
        <xref ref-type="bibr" rid="ref17">17</xref>
        ].
An example of coastal ecohydrology is the case of the Guadiana River estuary (Portugal), declared an
Ecohydrology Demonstration Site (EDS) by UNESCO in 2006 [
        <xref ref-type="bibr" rid="ref19">19</xref>
        ], where a dam was built to reduce
dependence on water from Spain and the irrigation of 22.3% in the agricultural area [
        <xref ref-type="bibr" rid="ref16 ref19">16, 19</xref>
        ]. Another
example is the Lacar Lake EDS project (Argentina), recognised in 2005, representing Latin America and
the Caribbean, by exhibiting the application of ecohydrological approaches in a basin [
        <xref ref-type="bibr" rid="ref15">15</xref>
        ]. In Ecuador
there are three EDS located in the Andean, coastal and island regions, being Palta Catacocha, Loja
(2018) the first EDS in Ecuador, considered as an artificial reservoir where rain and runof water are
used to recharge the associated aquifer [
        <xref ref-type="bibr" rid="ref16">16</xref>
        ], Pelican Bay, Galapagos (2019) stands out for its ecological
importance, and the management of water resources on Santa Cruz Island [
        <xref ref-type="bibr" rid="ref15">15</xref>
        ] and the most recent
Manglaralto, Santa Elena (2025) which is characterised by the river-aquifer system and its role in
supplying rural coastal communities [
        <xref ref-type="bibr" rid="ref20">20</xref>
        ].
      </p>
      <p>
        The province of Santa Elena (SEP) is a semi-arid area, whose freshwater distribution resources are
diferentiated according to geographical location and resources. For example, the southern zone of
the SEP is based on the Chongo-Colonche and Azúcar reservoirs of the Daule-Guayas River transfer
and the administration of the public company [
        <xref ref-type="bibr" rid="ref21 ref22">21, 22</xref>
        ]. The northern zone depends on coastal aquifers
and the management of community water organisations [
        <xref ref-type="bibr" rid="ref23">23</xref>
        ]. Manglaralto is north of the SEP and
has two types of climates characterised by the season of intense rains (February-April) and moderate
rains, "garuas" (July-December) [
        <xref ref-type="bibr" rid="ref24 ref25">24, 25</xref>
        ], with temperatures that fluctuate between 22 °C and 26 °C.
This commune is made up of an impermeable lithology favouring the accumulation of water, while
the permeable layer is composed of Quaternary alluvial deposits, where the inhabitants have taken
advantage of underground water resources through artisanal wells [
        <xref ref-type="bibr" rid="ref20">20</xref>
        ].
      </p>
      <p>
        The Manglaralto River-Aquifer System is a project declared as an EDS in March 2025, which stands
out for the influence of the "tape" (technical-artisanal dam) in the biota-hydrology processes of the
Manglaralto River sub-basin [
        <xref ref-type="bibr" rid="ref20">20</xref>
        ]. This hydraulic structure allows the retention of rainwater flows in
the seasonal riverbed, allowing infiltration into the soil and recharging the aquifer (sowing process),
to be later used through wells built by the community organisation Junta Administradora de Agua
Potable Regional Manglaralto (JAAPMAN) for controlled extraction in times of drought (harvesting
process) [
        <xref ref-type="bibr" rid="ref24">24</xref>
        ]. In this context, it is necessary to evaluate the tourist carrying capacity at demonstration
sites, such as the Manglaralto River-Aquifer System, to establish strategies according to sustainability
trends and water resource management. Coastal areas are places of connection between land and ocean,
where beach tourism is carried out to improve socioeconomic conditions. This anthropic activity poses
particular challenges in the degradation and loss of the marine-coastal ecosystem [
        <xref ref-type="bibr" rid="ref26 ref27">26, 27</xref>
        ]. Faced with
this problem, the following research questions are posed: What is the tourist carrying capacity of the
Manglaralto River-Aquifer System? What strategies are proposed to contribute to the ecohydrological,
hydrogeological and socioeconomic sustainability of the UNESCO ecohydrology demonstration site?
The study aims to evaluate the tourist carrying capacity in the Manglaralto River-Aquifer System,
through physical, ecological, and social indicators and the cause-efect methodology, to obtain the
minimum number of tourists necessary for the definition of sustainability strategies for water resources.
      </p>
    </sec>
    <sec id="sec-2">
      <title>2. Material and methods</title>
      <p>This study evaluates quantitative and qualitative techniques for analysing tourism carrying capacity
within the Manglaralto River-Aquifer system using environmental and social criteria. The process
involves three research phases: i) data collection and analysis, ii) assessment of tourism carrying
capacity, and iii) formulation of sustainability strategies (Figure 1).</p>
      <sec id="sec-2-1">
        <title>2.1. Phase I: Data collection and analysis</title>
        <p>
          The first phase consisted of the collection of primary and secondary information using various tools
for documentary and spatial analysis, field observations, and open interviews. Data on the number of
inhabitants were obtained by analysing population information from oficial sources, such as the census
conducted by the National Institute of Statistics and Census (INEC) [
          <xref ref-type="bibr" rid="ref28">28</xref>
          ]. Meanwhile, available visitor
records were provided by JAAPMAN, and the type of tourist, duration, and main activities carried out
during their stay were evaluated.
        </p>
        <p>Additionally, during this stage, accessibility, services, and tourism infrastructure were evaluated
through observation, field visits, open-ended interviews with the water board president and operators,
and the use of georeferenced tracking mobile devices (Geo Tracker version 5.4.1.4346). This application
allowed us to determine the distance and duration of the trip. The study area was delimited by a
topographic survey using diferential GPS (EGM 1996) and a drone (DJI Mavic Pro-2). The images
obtained were processed using Agisoft Metashape (version 1.8.4) and QGIS software to obtain Digital
Elevation Models and orthomosaics. This spatial information provided information on the area and
physical characteristics of the site.</p>
      </sec>
      <sec id="sec-2-2">
        <title>2.2. Phase II: Assessment of tourism carrying capacity</title>
        <p>
          Phase II evaluated tourist carrying capacity following the method proposed by Cifuentes [
          <xref ref-type="bibr" rid="ref29">29</xref>
          ], which is
used to estimate the maximum number of visitors a site can accommodate based on physical, social,
environmental, and administrative conditions [
          <xref ref-type="bibr" rid="ref30 ref31">30, 31</xref>
          ]. Three components were considered [
          <xref ref-type="bibr" rid="ref32">32</xref>
          ]:
• Physical Carrying Capacity (PCC): This is the limit of tourists that a site can receive in a specific
time and space.
• Carrying Capacity (RCC): This is composed of correction factors that directly and indirectly afect
the site.
        </p>
        <p>• Efective Carrying Capacity (ECC): This is the management or administrative capacity of the site.</p>
      </sec>
      <sec id="sec-2-3">
        <title>Physical carrying capacity (PCC)</title>
        <p>
          To calculate the physical carrying capacity of the SDE, the following parameters were determined:
the total area of the site, the number of daily hours it is open to the public, the average duration
of a tourist visit, and, in the case of the trail, the estimated time it takes for a visitor to complete
the route. These elements formed the basis for evaluating the physical carrying capacity of the site,
without considering additional factors such as vegetation, infrastructure, or ecological fragility. Physical
carrying capacity represents the maximum number of visitors a site can accommodate simultaneously
under ideal conditions without generating immediate negative impacts. This value was calculated using
the following equation 1, proposed by Cifuentes [
          <xref ref-type="bibr" rid="ref29">29</xref>
          ]:
 =


. 
(1)
        </p>
        <p>Where:  =Physical Carrying Capacity,  = total area of the site,  = area occupied by a
person,   = number of times the site can be visited.</p>
      </sec>
      <sec id="sec-2-4">
        <title>Real carrying capacity</title>
        <p>
          To determine the real capacity, it was necessary to decide on the correction factors according to Cifuentes
[
          <xref ref-type="bibr" rid="ref29">29</xref>
          ]. In this study, the factors were social (how many people enter), erodibility (the eroded area of the
site), rainfall per year, solar radiation per year, limiting days per year (for maintenance or nesting),
vegetation, and accessibility [
          <xref ref-type="bibr" rid="ref29">29</xref>
          ]. Subsequently, the actual load capacity of the site was calculated
according to Cifuentes’ equation 2:
        </p>
        <p>Where:  = Real Carrying capacity,   = Physical Carrying capacity,  , ... =
correction factors.</p>
      </sec>
      <sec id="sec-2-5">
        <title>Efective Carrying Capacity (ECC)</title>
        <p>Additionally, the ECC was determined by the product between the RCC and the handling capacity (HC),
which is presented in equation 3:</p>
        <p>Where:  = Carrying capacity,  = Real Carrying Capacity,  = Handling capacity
represented by the equation 4:</p>
        <p>=  * 
 =
 +   + 
3</p>
      </sec>
      <sec id="sec-2-6">
        <title>Cause-and-efect matrix</title>
        <p>
          The cause-efect matrix is a tool to identify the efect based on a set of qualitative characteristics to
measure the environmental impact [
          <xref ref-type="bibr" rid="ref33 ref34">33, 34</xref>
          ]. In this case, eight tourist activities were considered, and
the afected component, based on the magnitude, was observed. The characteristics to be considered
are listed in Table 1:
(2)
(3)
(4)
(5)
        </p>
        <p>=   *   * ( +  +  + )</p>
        <p>Where:   = Nature,   = Probability,  = Duration,  = Revesibility,  = Intensity,
 = Exclusivity.</p>
        <p>
          Table 2 establishes the qualitative and quantitative ranges used to evaluate the impact of tourism
activities. The evaluation considers criteria such as the magnitude range of the impact, which ranges
from -9 to +7, and the interpretation of the impact, ranging from high negative to high positive [
          <xref ref-type="bibr" rid="ref34">34</xref>
          ].
        </p>
      </sec>
      <sec id="sec-2-7">
        <title>2.3. Phase III: Strategies for sustainability</title>
        <p>
          To ensure efective and participatory planning, the creation of a multidisciplinary panel for the
formulation of an environmental strategy was proposed. The panel comprised 13 members selected
based on rigorous criteria, such as technical experience in hydrology, ecology, geology, and tourism,
as well as community leadership, institutional representation, and local cultural knowledge from the
six communities benefiting from the JAAPMAN service. The composition of the panel included the
president of the water board, two technical operators of the system, four geological engineers and
researchers related to the geosciences, an environmental engineer, a tourism graduate, a civil engineer,
and three members of the financial administration of JAAPMAN. The panel was held in person through
the workshop ’Main Challenges in the Management of the Demonstrative Ecohydrology Site’, held on
January 1, 2025, where three rounds were sequentially developed, aimed at creating environmental
strategies for the site:
1. Conceptualization stage: Where the concept of a ’Demonstration Site’ was defined, integrating
ecological, technical, and community impacts.
2. Participatory diagnosis stage: Strengths, Weaknesses, Opportunities, and Threats (SWOT)
analysis tools [
          <xref ref-type="bibr" rid="ref21">21</xref>
          ] were applied to evaluate the Demonstration Site project to inform sustainable
decision-making. This analysis indicates the strengths, opportunities, weaknesses, and threats of
the site from the perspective of the representatives of each expert panel. They contributed ideas
based on their local experience and shared their vision regarding the challenges and potential
opportunities to enhance the site.
3. Stage of strategy planning and socialization: Strategies were implemented to manage
sustainable tourism, aimed at reducing the impact on the ecosystem and improving the tourist experience
in the DSE. These strategies were shared among specialists and community representatives. And
it was carried out by applying the crossing of the aspects established in the SWOT analysis.
        </p>
      </sec>
    </sec>
    <sec id="sec-3">
      <title>3. Results and discussion</title>
      <sec id="sec-3-1">
        <title>3.1. Baseline diagnosis and spatial characterization of the demonstration site of the</title>
      </sec>
      <sec id="sec-3-2">
        <title>Manglaralto-Aquifer River</title>
        <p>The DSE has an area of 1891.55 m2, of which 809 m2 is enabled for tourism activities (figure 2), while the
remaining area is covered by vegetation. The walking tour of the site takes approximately 20 minutes,
starting at the pumping station, where an introduction to the local water catchment system is provided.
Next, the visit proceeds to Well 1, where historical and technical information about the construction of
the well is provided, highlighting the role of international cooperation projects between ESPOL and the
International Atomic Energy Agency (IAEA). At this point, monthly monitoring of physicochemical
analyses and static/dynamic water levels (Figure 3a) is regularly conducted, allowing for the assessment
of its quality and hydrogeological characteristics. Well 2 serves the purpose of monitoring the static
level of the aquifer, providing essential information for understanding the behaviour of the underground
system and the state of the water table at diferent times of the year.</p>
        <p>Continuing with the tour, we reached the halfway point of the journey, where the upstream
environment of the Manglaralto River can be observed (Figure 3b). In front, there is an observation area for the
endemic species of the region (Figures 4a and 4b), which enhances the ecological experience of the trail.
Moving along the route, the journey concluded at the ’tape’ dam, a place used by residents for artisanal
shrimp fishing and as a water recreation area. On warm afternoons, this site becomes a gathering place
where dozens of families enjoy the natural scenery and the freshwater of the Manglaralto River (Figure
4c) for almost three hours. Furthermore, the ’tape’ is used like a didactic example of the technique for
sowing and harvesting water that facilitates the recharge of the coastal aquifer.
(a) Monitoring activity of
levels and physicochemical
parameters of wells.</p>
        <p>(b) View of the upper part of the Manglaralto River.</p>
        <p>Regarding the site’s basic infrastructure, it is enclosed by fencing but lacks essential amenities such
as signage, paved pathways, sanitary facilities, food courts, information kiosks, and waste disposal
services. From an environmental management perspective, the only existing measure is a warning sign
indicating deep waters, which is poorly visible due to surrounding vegetation. From a tourist service
perspective, there are no designated guides available during visiting hours to monitor visitor numbers
or to tour the characteristics of the Manglaralto River–Aquifer System.</p>
      </sec>
      <sec id="sec-3-3">
        <title>3.2. Assessment of environmental impacts and tourism carrying capacity of the site</title>
        <p>The FCC of the DSE is 2523 visitors per day, considering the entire site area. To obtain the RCC,
correction factors were considered, and a total of 698 visitors per day was obtained. In the ECC, a 0.33%
(a) Golden-toed Heron (Egretta thula)</p>
        <p>(b) Blue Heron (Ardea herodias).</p>
        <p>
          (c) Panoramic view of the ’tape’ dam
was obtained considering the handling capacity at the site, with a daily tourist load capacity of 24 people
without causing ecological collapse. A similar case study is the Complejo Educativo Ambiental Naciente
Arriaz (CEANA) in Costa Rica, which encompasses 4.31 hectares of forest protecting a spring that is
used for supplying drinking water to the communities of “Taras”, “La Lima”, and part of “La Fátima”,
where ecotourism activities are conducted. At CEANA, it was established that the three trails should be
used by an average of 26 people per day to maintain the ecological balance of the site in relation to the
spring season. However, before 2019, this spring was afected by vandalism and pollution, which led to
the creation of a fence around it to ensure the sustainability of the site [
          <xref ref-type="bibr" rid="ref35">35</xref>
          ]. Even in UNESCO World
Heritage sites like Angkor, it has been shown that a 6% increase in annual visits can cause impacts on the
unique elements of this type of site, including temples and heritage monuments, as well as generating
impacts on freshwater sources due to excessive tourism, such as the extraction of more than 27,900 m³
of groundwater [
          <xref ref-type="bibr" rid="ref36">36</xref>
          ]. According to scientific literature [
          <xref ref-type="bibr" rid="ref37 ref38">37, 38</xref>
          ] sustained tourist pressure on ecosystems
can lead to gradual degradation of their natural and cultural attributes, ultimately rendering the site
unsuitable for visitation. In the context of UNESCO demonstration sites, where the dual objective is to
promote sustainable use while conserving ecological integrity, calculating the tourist carrying capacity
becomes a key management tool.
        </p>
        <p>For the analysis of the cause-efect matrix, tourist activity (cause) and the afected component (efect)
were considered. Recreational activities, such as walking on unpaved trails, bird watching, artisanal
ifshing, excessive use of the dam, presence of garbage in the dam, and vehicle frequency, were found
to have negative impacts, with magnitudes between -9 and -3.5 (Table 3). Meanwhile, activities that
integrate socio-educational values had positive impacts with magnitudes of +4 and +5.</p>
      </sec>
      <sec id="sec-3-4">
        <title>3.3. Strategic guidelines for sustainable tourism in a Manglaralto River-Aquifer demonstration site</title>
        <p>• Basic infrastructure and tourism use planning:(strategies derived from WO+WA): This
includes the development of minimum services (bathrooms, signage, resting areas), site zoning,
control of tourist carrying capacity, and implementation of a tourism management and monitoring
plan.
• Environmental education and strengthening the scientific value of the site: (strategies
derived from SO+WO): designing an eco-hydrological and geotourism interpretive trail, generating
workshops, guided tours, informational panels, and training local guides. In addition to promoting
technical, scientific, and educational tourism at the local level.
• Conservation of biodiversity and ecological restoration: (strategies derived from OT+WO):
implementation of riparian vegetation restoration programs, control of invasive species and
habitat recovery, development of a biological inventory program, and participatory monitoring
of flora and fauna in partnership with academia-research.
• Participatory management and institutional articulation: (strategies derived from
SO+WT+OT): consolidation of a local management unit of the site (JAAPMAN+allies),
strengthening of alliances with universities, tourism stakeholders, and cooperation agencies, and
development of promotional campaigns for the site in networks of geosites and protected areas at the
national and international levels.</p>
      </sec>
    </sec>
    <sec id="sec-4">
      <title>4. Conclusions</title>
      <p>The study established a tourist load of a minimum of 24 tourists per day, keeping the negative
impacts of tourism under control. Through the cause-efect matrix, the most sensitive environmental
components were identified. For example, excessive use of the dam for recreational activities could
generate significant wear, classified as a high negative impact with a magnitude of -9. In contrast, the
implementation of proper signage would contribute to the appreciation of the landscape, classified as a
medium positive impact with a magnitude of +5.</p>
      <p>Strategies were proposed in the structural area, such as the design of a geotouristic interpretative trail,
development of minimum services (bathrooms, signage, resting areas), site zoning, and implementation
of a tourism management and monitoring plan. From an educational perspective, the creation of
a community centre for environmental education and technical tourism, as well as workshops for
community training in environmental guidance and participatory monitoring, was proposed. This study
used quantitative approaches based on physical, social, environmental, and administrative conditions,
as well as the perceptions of key stakeholders and experts. State space model studies are necessary to
quantify carrying capacity from the perspective of ecological eficiency for sustainable and dynamic
management of carrying capacity at sites of coastal ecohydrological importance, such as the case study.</p>
    </sec>
    <sec id="sec-5">
      <title>Acknowledgements</title>
      <p>The authors are grateful for the support provided through the research project "Registry of
Geological Sites of Interest in Ecuador for Sustainable Development Strategies" (CIPAT-004-2024), which
strengthened the tourism, ecological, and hydrogeological components of this case study. They also
acknowledge the support of the community engagement project "Sowing Harvesting and Reusing
Water for Sustainability (Phase II)" (PG13-PY25-07) and JAAPMAN, whose key stakeholders actively
participated in the various stages of the research process.</p>
    </sec>
    <sec id="sec-6">
      <title>Declaration on Generative AI</title>
      <p>The authors have not employed any Generative AI tools.</p>
    </sec>
  </body>
  <back>
    <ref-list>
      <ref id="ref1">
        <mixed-citation>
          [1]
          <string-name>
            <given-names>C. M.</given-names>
            <surname>Hall</surname>
          </string-name>
          ,
          <article-title>Trends in ocean and coastal tourism: the end of the last frontier?</article-title>
          ,
          <source>Ocean &amp; coastal management 44</source>
          (
          <year>2001</year>
          )
          <fpage>601</fpage>
          -
          <lpage>618</lpage>
          .
        </mixed-citation>
      </ref>
      <ref id="ref2">
        <mixed-citation>
          [2]
          <string-name>
            <given-names>S.</given-names>
            <surname>Hynes</surname>
          </string-name>
          ,
          <string-name>
            <given-names>M.</given-names>
            <surname>Cawley</surname>
          </string-name>
          ,
          <string-name>
            <given-names>J.</given-names>
            <surname>Deely</surname>
          </string-name>
          ,
          <string-name>
            <given-names>D.</given-names>
            <surname>Norton</surname>
          </string-name>
          ,
          <article-title>Alternative approaches to measuring the value of tourism in marine and coastal areas in ocean economy accounting</article-title>
          ,
          <source>Marine Policy</source>
          <volume>168</volume>
          (
          <year>2024</year>
          )
          <fpage>106299</fpage>
          .
        </mixed-citation>
      </ref>
      <ref id="ref3">
        <mixed-citation>
          [3] R. de la Cruz Modino,
          <article-title>Turismo marino</article-title>
          . desarrollo, impactos y gestión.,
          <source>PASOS Revista de Turismo y Patrimonio Cultural</source>
          <volume>4</volume>
          (
          <year>2006</year>
          )
          <fpage>117</fpage>
          -
          <lpage>122</lpage>
          .
        </mixed-citation>
      </ref>
      <ref id="ref4">
        <mixed-citation>
          [4]
          <string-name>
            <given-names>G.</given-names>
            <surname>Mendoza-González</surname>
          </string-name>
          ,
          <string-name>
            <given-names>M. L.</given-names>
            <surname>Martínez</surname>
          </string-name>
          ,
          <string-name>
            <given-names>R.</given-names>
            <surname>Guevara</surname>
          </string-name>
          ,
          <string-name>
            <given-names>O.</given-names>
            <surname>Pérez-Maqueo</surname>
          </string-name>
          ,
          <string-name>
            <given-names>M. C.</given-names>
            <surname>Garza-Lagler</surname>
          </string-name>
          ,
          <string-name>
            <given-names>A.</given-names>
            <surname>Howard</surname>
          </string-name>
          ,
          <article-title>Towards a sustainable sun, sea, and sand tourism: The value of ocean view and proximity to the coast</article-title>
          ,
          <source>Sustainability</source>
          <volume>10</volume>
          (
          <year>2018</year>
          )
          <fpage>1012</fpage>
          .
        </mixed-citation>
      </ref>
      <ref id="ref5">
        <mixed-citation>
          [5]
          <string-name>
            <given-names>L.</given-names>
            <surname>Fanini</surname>
          </string-name>
          ,
          <string-name>
            <given-names>L. L.</given-names>
            <surname>Costa</surname>
          </string-name>
          ,
          <string-name>
            <given-names>I. R.</given-names>
            <surname>Zalmon</surname>
          </string-name>
          ,
          <string-name>
            <given-names>M.</given-names>
            <surname>Riechers</surname>
          </string-name>
          ,
          <article-title>Social and ecological elements for a perspective approach to citizen science on the beach</article-title>
          ,
          <source>Frontiers in Ecology and Evolution</source>
          <volume>9</volume>
          (
          <year>2021</year>
          )
          <fpage>694487</fpage>
          .
        </mixed-citation>
      </ref>
      <ref id="ref6">
        <mixed-citation>
          [6]
          <string-name>
            <given-names>E. P.</given-names>
            <surname>Odum</surname>
          </string-name>
          , et al., Fundamentos de ecologia,
          <year>1953</year>
          .
        </mixed-citation>
      </ref>
      <ref id="ref7">
        <mixed-citation>
          [7]
          <string-name>
            <given-names>E. P.</given-names>
            <surname>Odum</surname>
          </string-name>
          , et al., Fundamentos de ecologia,
          <year>2006</year>
          .
        </mixed-citation>
      </ref>
      <ref id="ref8">
        <mixed-citation>
          [8]
          <string-name>
            <given-names>E.</given-names>
            <surname>Odum</surname>
          </string-name>
          ,
          <article-title>La ecología y nuestros sistemas de soporte vital en peligro</article-title>
          , Editorial Sinauer Associates (
          <year>1993</year>
          ).
        </mixed-citation>
      </ref>
      <ref id="ref9">
        <mixed-citation>
          [9]
          <string-name>
            <surname>C.</surname>
          </string-name>
          <article-title>A</article-title>
          . T. de Investigación y Enseñanza. Catie,
          <string-name>
            <given-names>M.</given-names>
            <surname>Cifuentes</surname>
          </string-name>
          ,
          <string-name>
            <given-names>F.</given-names>
            <surname>Barroso</surname>
          </string-name>
          , W. Alpizar, Capacidad de carga turística de la Reserva Biológica Carara, Centro Agronomico Tropical de Investigación y Enseñanza,
          <year>1990</year>
          .
        </mixed-citation>
      </ref>
      <ref id="ref10">
        <mixed-citation>
          [10]
          <string-name>
            <given-names>O.</given-names>
            <surname>Tokarchuk</surname>
          </string-name>
          ,
          <string-name>
            <given-names>R.</given-names>
            <surname>Gabriele</surname>
          </string-name>
          ,
          <string-name>
            <given-names>O.</given-names>
            <surname>Maurer</surname>
          </string-name>
          ,
          <article-title>Estimating tourism social carrying capacity</article-title>
          ,
          <source>Annals of Tourism Research</source>
          <volume>86</volume>
          (
          <year>2021</year>
          )
          <fpage>102971</fpage>
          .
        </mixed-citation>
      </ref>
      <ref id="ref11">
        <mixed-citation>
          [11]
          <string-name>
            <surname>A. M. O'Reilly</surname>
          </string-name>
          ,
          <article-title>Tourism carrying capacity: Concept and issues</article-title>
          ,
          <source>Tourism management 7</source>
          (
          <year>1986</year>
          )
          <fpage>254</fpage>
          -
          <lpage>258</lpage>
          .
        </mixed-citation>
      </ref>
      <ref id="ref12">
        <mixed-citation>
          [12]
          <string-name>
            <surname>J. R. de Souza</surname>
            <given-names>Filho</given-names>
          </string-name>
          ,
          <string-name>
            <given-names>R. C.</given-names>
            <surname>Santos</surname>
          </string-name>
          ,
          <string-name>
            <given-names>I. R.</given-names>
            <surname>Silva</surname>
          </string-name>
          ,
          <string-name>
            <surname>C. I. Ellif</surname>
          </string-name>
          ,
          <article-title>Evaluation of recreational quality, carrying capacity and ecosystem services supplied by sandy beaches of the municipality of camaçari, northern coast of bahia, brazil</article-title>
          ,
          <source>Journal of Coastal Research</source>
          (
          <year>2014</year>
          )
          <fpage>527</fpage>
          -
          <lpage>532</lpage>
          .
        </mixed-citation>
      </ref>
      <ref id="ref13">
        <mixed-citation>
          [13]
          <string-name>
            <given-names>E. J.</given-names>
            <surname>Wilkins</surname>
          </string-name>
          ,
          <string-name>
            <given-names>Y.</given-names>
            <surname>Chikamoto</surname>
          </string-name>
          ,
          <string-name>
            <given-names>A. B.</given-names>
            <surname>Miller</surname>
          </string-name>
          ,
          <string-name>
            <given-names>J. W.</given-names>
            <surname>Smith</surname>
          </string-name>
          ,
          <article-title>Climate change and the demand for recreational ecosystem services on public lands in the continental united states</article-title>
          ,
          <source>Global Environmental Change</source>
          <volume>70</volume>
          (
          <year>2021</year>
          )
          <fpage>102365</fpage>
          .
        </mixed-citation>
      </ref>
      <ref id="ref14">
        <mixed-citation>
          [14]
          <article-title>UNESCO</article-title>
          and
          <string-name>
            <surname>IHP</surname>
          </string-name>
          , Demonstrative sites ecohydrology,
          <year>2025</year>
          .
        </mixed-citation>
      </ref>
      <ref id="ref15">
        <mixed-citation>
          [15]
          <string-name>
            <given-names>M.</given-names>
            <surname>Gaviño Novillo</surname>
          </string-name>
          ,
          <string-name>
            <given-names>M.</given-names>
            <surname>Doria</surname>
          </string-name>
          , et al.,
          <article-title>Sitios demostrativos de ecohidrología en américa latina y el caribe: pasado</article-title>
          , presente y futuro,
          <year>2024</year>
          .
        </mixed-citation>
      </ref>
      <ref id="ref16">
        <mixed-citation>
          [16]
          <string-name>
            <given-names>M.</given-names>
            <surname>Albarracín</surname>
          </string-name>
          , G. Ramón,
          <string-name>
            <given-names>J.</given-names>
            <surname>González</surname>
          </string-name>
          ,
          <string-name>
            <given-names>C.</given-names>
            <surname>Iñiguez-Armijos</surname>
          </string-name>
          ,
          <string-name>
            <given-names>T.</given-names>
            <surname>Zakaluk</surname>
          </string-name>
          ,
          <string-name>
            <given-names>S.</given-names>
            <surname>Martos-Rosillo</surname>
          </string-name>
          ,
          <article-title>The ecohydrological approach in water sowing and harvesting systems: The case of the paltas catacocha ecohydrology demonstration site, ecuador</article-title>
          ,
          <source>Ecohydrology &amp; Hydrobiology</source>
          <volume>21</volume>
          (
          <year>2021</year>
          )
          <fpage>454</fpage>
          -
          <lpage>466</lpage>
          .
        </mixed-citation>
      </ref>
      <ref id="ref17">
        <mixed-citation>
          [17]
          <string-name>
            <surname>UNESCO</surname>
          </string-name>
          ,
          <article-title>La ecohidrología: aportes concretos al desarrollo sostenible</article-title>
          ,
          <year>2025</year>
          .
        </mixed-citation>
      </ref>
      <ref id="ref18">
        <mixed-citation>
          [18]
          <string-name>
            <given-names>S.</given-names>
            <surname>Ayari</surname>
          </string-name>
          ,
          <string-name>
            <given-names>H.</given-names>
            <surname>Gabtni</surname>
          </string-name>
          , L. Zouhri,
          <article-title>Imaging seawater flow in porous media under climate change water stress using geophysics: a case study of menzel horr coastline in northeastern tunisia, mediterranean basin</article-title>
          ,
          <source>Environmental Earth Sciences</source>
          <volume>82</volume>
          (
          <year>2023</year>
          )
          <fpage>499</fpage>
          .
        </mixed-citation>
      </ref>
      <ref id="ref19">
        <mixed-citation>
          [19]
          <string-name>
            <given-names>L.</given-names>
            <surname>Chícharo</surname>
          </string-name>
          ,
          <string-name>
            <given-names>J.</given-names>
            <surname>Gaona</surname>
          </string-name>
          ,
          <string-name>
            <given-names>P.</given-names>
            <surname>Sousa</surname>
          </string-name>
          ,
          <string-name>
            <given-names>G.</given-names>
            <surname>Arduino</surname>
          </string-name>
          ,
          <article-title>A methodology to evaluate the implementation of ecohydrology solutions at the ecohydrology demonstration sites: The guadiana case study</article-title>
          ,
          <source>Ecohydrology &amp; Hydrobiology</source>
          <volume>22</volume>
          (
          <year>2022</year>
          )
          <fpage>391</fpage>
          -
          <lpage>399</lpage>
          .
        </mixed-citation>
      </ref>
      <ref id="ref20">
        <mixed-citation>
          [20]
          <string-name>
            <surname>UNESCO</surname>
          </string-name>
          , Manglaralto, nuevo sitio demostrativo de ecohidrología de la unesco en ecuador,
          <year>2025</year>
          .
        </mixed-citation>
      </ref>
      <ref id="ref21">
        <mixed-citation>
          [21]
          <string-name>
            <given-names>G.</given-names>
            <surname>Herrera-Franco</surname>
          </string-name>
          ,
          <string-name>
            <given-names>F.</given-names>
            <surname>Morante-Carballo</surname>
          </string-name>
          ,
          <string-name>
            <given-names>L.</given-names>
            <surname>Bravo-Montero</surname>
          </string-name>
          ,
          <string-name>
            <given-names>J.</given-names>
            <surname>Valencia-Robles</surname>
          </string-name>
          ,
          <string-name>
            <given-names>M.</given-names>
            <surname>Aguilar-Aguilar</surname>
          </string-name>
          ,
          <string-name>
            <given-names>S.</given-names>
            <surname>Martos-Rosillo</surname>
          </string-name>
          ,
          <string-name>
            <given-names>P.</given-names>
            <surname>Carrión-Mero</surname>
          </string-name>
          ,
          <article-title>Water sowing and harvesting (ws&amp;h) for sustainable management in ecuador: A review</article-title>
          ,
          <source>Heritage</source>
          <volume>7</volume>
          (
          <year>2024</year>
          )
          <fpage>3696</fpage>
          -
          <lpage>3718</lpage>
          .
        </mixed-citation>
      </ref>
      <ref id="ref22">
        <mixed-citation>
          [22]
          <string-name>
            <given-names>C.</given-names>
            <surname>Cornejo</surname>
          </string-name>
          ,
          <string-name>
            <given-names>D. Z.</given-names>
            <surname>Haman</surname>
          </string-name>
          ,
          <string-name>
            <given-names>R. L.</given-names>
            <surname>Espinel</surname>
          </string-name>
          ,
          <string-name>
            <given-names>J.</given-names>
            <surname>Jordan</surname>
          </string-name>
          ,
          <article-title>Irrigation potential of the trasvase system (santa elena peninsula, guayas</article-title>
          , ecuador),
          <source>Journal of irrigation and drainage engineering 132</source>
          (
          <year>2006</year>
          )
          <fpage>453</fpage>
          -
          <lpage>462</lpage>
          .
        </mixed-citation>
      </ref>
      <ref id="ref23">
        <mixed-citation>
          [23]
          <string-name>
            <given-names>F.</given-names>
            <surname>Morante-Carballo</surname>
          </string-name>
          ,
          <string-name>
            <given-names>J.</given-names>
            <surname>Briones-Bitar</surname>
          </string-name>
          ,
          <string-name>
            <given-names>F.</given-names>
            <surname>Montalván</surname>
          </string-name>
          ,
          <string-name>
            <given-names>A.</given-names>
            <surname>Alencastro-Segura</surname>
          </string-name>
          ,
          <string-name>
            <given-names>M. A.</given-names>
            <surname>Chávez-Moncayo</surname>
          </string-name>
          ,
          <string-name>
            <given-names>P.</given-names>
            <surname>Carrión-Mero</surname>
          </string-name>
          ,
          <article-title>Proposal of an alluvial dike as a nature-based solution for sustainable water management in coastal areas</article-title>
          ,
          <source>Results in Engineering</source>
          <volume>23</volume>
          (
          <year>2024</year>
          )
          <fpage>102599</fpage>
          .
        </mixed-citation>
      </ref>
      <ref id="ref24">
        <mixed-citation>
          [24]
          <string-name>
            <surname>GADM</surname>
          </string-name>
          , Clima de la parroquia manglaralto,
          <year>2025</year>
          .
        </mixed-citation>
      </ref>
      <ref id="ref25">
        <mixed-citation>
          [25]
          <string-name>
            <given-names>M.</given-names>
            <surname>Leon</surname>
          </string-name>
          , G. Cornejo,
          <string-name>
            <given-names>M.</given-names>
            <surname>Calderón</surname>
          </string-name>
          ,
          <string-name>
            <given-names>E.</given-names>
            <surname>González-Carrión</surname>
          </string-name>
          ,
          <string-name>
            <given-names>H.</given-names>
            <surname>Florez</surname>
          </string-name>
          ,
          <article-title>Efect of deforestation on climate change: A co-integration and causality approach with time series</article-title>
          ,
          <source>Sustainability</source>
          <volume>14</volume>
          (
          <year>2022</year>
          )
          <fpage>1</fpage>
          -
          <lpage>14</lpage>
          . doi:
          <volume>10</volume>
          .3390/su141811303.
        </mixed-citation>
      </ref>
      <ref id="ref26">
        <mixed-citation>
          [26]
          <string-name>
            <given-names>G.</given-names>
            <surname>Skiniti</surname>
          </string-name>
          ,
          <string-name>
            <given-names>M.</given-names>
            <surname>Lilli</surname>
          </string-name>
          ,
          <string-name>
            <given-names>N.</given-names>
            <surname>Skarakis</surname>
          </string-name>
          ,
          <string-name>
            <given-names>S.</given-names>
            <surname>Tournaki</surname>
          </string-name>
          ,
          <string-name>
            <given-names>N.</given-names>
            <surname>Nikolaidis</surname>
          </string-name>
          ,
          <string-name>
            <given-names>T.</given-names>
            <surname>Tsoutsos</surname>
          </string-name>
          ,
          <article-title>A holistic approach for tourism carrying capacity estimation in sensitive ecological areas</article-title>
          ,
          <source>Environment, Development and Sustainability</source>
          <volume>26</volume>
          (
          <year>2024</year>
          )
          <fpage>31971</fpage>
          -
          <lpage>31995</lpage>
          .
        </mixed-citation>
      </ref>
      <ref id="ref27">
        <mixed-citation>
          [27]
          <string-name>
            <given-names>B.</given-names>
            <surname>Dibba</surname>
          </string-name>
          ,
          <string-name>
            <given-names>S.</given-names>
            <surname>Yafa</surname>
          </string-name>
          ,
          <string-name>
            <given-names>M.</given-names>
            <surname>Sawaneh</surname>
          </string-name>
          , W. Adzawla,
          <article-title>Land cover transformation and population growth: Impacts on coastal environment of the gambia (</article-title>
          <year>1990</year>
          -2020),
          <source>Sustainability</source>
          <volume>17</volume>
          (
          <year>2025</year>
          )
          <year>1853</year>
          .
        </mixed-citation>
      </ref>
      <ref id="ref28">
        <mixed-citation>
          [28]
          <string-name>
            <surname>INEC</surname>
          </string-name>
          , Censo ecuador cuenta conmigo,
          <year>2025</year>
          .
        </mixed-citation>
      </ref>
      <ref id="ref29">
        <mixed-citation>
          [29]
          <string-name>
            <given-names>M.</given-names>
            <surname>Cifuentes</surname>
          </string-name>
          , Determinación de capacidad de carga turística enáreas protegidas,
          <volume>194</volume>
          ,
          <string-name>
            <surname>Bib</surname>
          </string-name>
          .
          <source>Orton IICA/CATIE</source>
          ,
          <year>1992</year>
          .
        </mixed-citation>
      </ref>
      <ref id="ref30">
        <mixed-citation>
          [30]
          <string-name>
            <given-names>L. L.</given-names>
            <surname>Diniz</surname>
          </string-name>
          ,
          <string-name>
            <given-names>P. M.</given-names>
            <surname>Machado</surname>
          </string-name>
          , A. B. do
          <string-name>
            <surname>Nascimento</surname>
            ,
            <given-names>L. L.</given-names>
          </string-name>
          <string-name>
            <surname>Costa</surname>
          </string-name>
          , I. D. da Costa,
          <string-name>
            <surname>C. A. M. M. Cordeiro</surname>
            ,
            <given-names>I. R.</given-names>
          </string-name>
          <string-name>
            <surname>Zalmon</surname>
          </string-name>
          ,
          <article-title>Evaluation of tourist carrying capacity to support recreational beaches management</article-title>
          ,
          <source>Ocean &amp; Coastal Management</source>
          <volume>249</volume>
          (
          <year>2024</year>
          )
          <fpage>107022</fpage>
          .
        </mixed-citation>
      </ref>
      <ref id="ref31">
        <mixed-citation>
          [31]
          <string-name>
            <surname>I. Dias e Cordeiro</surname>
            ,
            <given-names>N.</given-names>
          </string-name>
          <string-name>
            <surname>Körössy</surname>
            ,
            <given-names>V. S.</given-names>
          </string-name>
          <string-name>
            <surname>Fragoso</surname>
            <given-names>Selva</given-names>
          </string-name>
          , Determinación de la capacidad de carga turística: El caso de playa de tamandaré-pernambuco-brasil,
          <source>Estudios y perspectivas en turismo 21</source>
          (
          <year>2012</year>
          )
          <fpage>0</fpage>
          -
          <lpage>0</lpage>
          .
        </mixed-citation>
      </ref>
      <ref id="ref32">
        <mixed-citation>
          [32]
          <string-name>
            <given-names>M.</given-names>
            <surname>Delgado</surname>
          </string-name>
          ,
          <article-title>Análise da metodologia criada por miguel cifuentes referente à capacidade de carga turística</article-title>
          ,
          <source>Revista Turismo em Análise</source>
          <volume>18</volume>
          (
          <year>2007</year>
          )
          <fpage>73</fpage>
          -
          <lpage>93</lpage>
          .
        </mixed-citation>
      </ref>
      <ref id="ref33">
        <mixed-citation>
          [33]
          <string-name>
            <given-names>P.</given-names>
            <surname>Carrion-Mero</surname>
          </string-name>
          ,
          <string-name>
            <given-names>L.</given-names>
            <surname>Soto-Navarrete</surname>
          </string-name>
          ,
          <string-name>
            <given-names>B.</given-names>
            <surname>Apolo-Masache</surname>
          </string-name>
          ,
          <string-name>
            <given-names>J.</given-names>
            <surname>Mata-Perello</surname>
          </string-name>
          ,
          <string-name>
            <given-names>G.</given-names>
            <surname>Herrera-Franco</surname>
          </string-name>
          ,
          <string-name>
            <given-names>J.</given-names>
            <surname>Briones-Bitar</surname>
          </string-name>
          ,
          <article-title>Environmental assessment and tourism carrying capacity in geosites of the ruta del oro geopark project</article-title>
          ,
          <source>Geoheritage</source>
          <volume>17</volume>
          (
          <year>2025</year>
          )
          <fpage>1</fpage>
          -
          <lpage>22</lpage>
          .
        </mixed-citation>
      </ref>
      <ref id="ref34">
        <mixed-citation>
          [34]
          <string-name>
            <given-names>A.</given-names>
            <surname>Garmendia</surname>
          </string-name>
          ,
          <string-name>
            <given-names>C.</given-names>
            <surname>Crespo</surname>
          </string-name>
          ,
          <string-name>
            <given-names>A.</given-names>
            <surname>Salvador</surname>
          </string-name>
          , Evaluación de impacto ambiental,
          <year>2005</year>
          .
        </mixed-citation>
      </ref>
      <ref id="ref35">
        <mixed-citation>
          [35]
          <string-name>
            <given-names>L.</given-names>
            <surname>Castro-Barrantes</surname>
          </string-name>
          ,
          <article-title>Capacidad de carga turística y zonificación del ceana, con miras al manejo de visitantes y protección del recurso hídrico en cartago, costa rica</article-title>
          ,
          <source>Revista Forestal Mesoamericana Kurú</source>
          <volume>17</volume>
          (
          <year>2020</year>
          )
          <fpage>47</fpage>
          -
          <lpage>57</lpage>
          .
        </mixed-citation>
      </ref>
      <ref id="ref36">
        <mixed-citation>
          [36]
          <string-name>
            <given-names>F.</given-names>
            <surname>Chen</surname>
          </string-name>
          ,
          <string-name>
            <given-names>H.</given-names>
            <surname>Guo</surname>
          </string-name>
          ,
          <string-name>
            <given-names>N.</given-names>
            <surname>Ishwaran</surname>
          </string-name>
          ,
          <string-name>
            <given-names>J.</given-names>
            <surname>Liu</surname>
          </string-name>
          ,
          <string-name>
            <given-names>X.</given-names>
            <surname>Wang</surname>
          </string-name>
          ,
          <string-name>
            <given-names>W.</given-names>
            <surname>Zhou</surname>
          </string-name>
          ,
          <string-name>
            <given-names>P.</given-names>
            <surname>Tang</surname>
          </string-name>
          ,
          <article-title>Understanding the relationship between the water crisis and sustainability of the angkor world heritage site</article-title>
          ,
          <source>Remote Sensing of Environment</source>
          <volume>232</volume>
          (
          <year>2019</year>
          )
          <fpage>111293</fpage>
          .
        </mixed-citation>
      </ref>
      <ref id="ref37">
        <mixed-citation>
          [37]
          <string-name>
            <given-names>R.</given-names>
            <surname>Butler</surname>
          </string-name>
          ,
          <string-name>
            <given-names>J. F.</given-names>
            <surname>Vera-Rebollo</surname>
          </string-name>
          ,
          <string-name>
            <given-names>I.</given-names>
            <surname>Rodriguez-Sanchez</surname>
          </string-name>
          , et al.,
          <article-title>Mature tourist destinations: can we recapture and retain the magic</article-title>
          ?,
          <year>2012</year>
          .
        </mixed-citation>
      </ref>
      <ref id="ref38">
        <mixed-citation>
          [38]
          <string-name>
            <given-names>P. A.</given-names>
            <surname>Aguilar Calderón</surname>
          </string-name>
          ,
          <string-name>
            <given-names>B. A.</given-names>
            <surname>López-Chávez</surname>
          </string-name>
          ,
          <string-name>
            <given-names>N. I. Peinado</given-names>
            <surname>Osuna</surname>
          </string-name>
          ,
          <string-name>
            <given-names>E. O. Burgueño</given-names>
            <surname>Sánchez</surname>
          </string-name>
          ,
          <string-name>
            <given-names>J. A.</given-names>
            <surname>Aguilar-Calderón</surname>
          </string-name>
          ,
          <string-name>
            <given-names>J. A.</given-names>
            <surname>Somoza</surname>
          </string-name>
          <string-name>
            <surname>Ríos</surname>
          </string-name>
          ,
          <article-title>Tourism carrying capacity in coastal destinations: An assessment in mazatlán, mexico</article-title>
          ,
          <source>Sustainability</source>
          <volume>17</volume>
          (
          <year>2025</year>
          )
          <fpage>6344</fpage>
          .
        </mixed-citation>
      </ref>
    </ref-list>
  </back>
</article>